Saturday, July 20, 2019

Enumerating California Needlegrasses (the tribe Stipeae)


Peterson PM, Romaschenko K, Soreng RJ, Reyna JV (2019) A key to the North American genera of Stipeae (Poaceae, Pooideae) with descriptions and taxonomic names for species of Eriocoma, Neotrinia, Oloptum, and five new genera: Barkworthia, ×Eriosella, Pseudoeriocoma, Ptilagrostiella, and Thorneochloa. PhytoKeys 126: 89–125. https://doi. org/10.3897/phytokeys.126.34096

California has by my count 42 taxa of needlegrasses: the Tribe Stipeae Dumort.  In the Jepson Manual, a broad (and overly simplified) generic concept was used: a single genus Stipa L.  Robustly supported phylogenetic and biogeographic data now indicates that a number of lineages qualify as genera, based on globally coherent viewpoint (yes, in Jepson-ease its so easy to just blurt out Stipa! and remain in ignorance).  Botanists struggle with the concept of a genus, and are, when confused, inclined to combine lineages and they don’t know when to stop.  This results inappropriate and ill-informative treatments: one could recognize only Triticum L. for all species within the Triticeae, the cereal grasses, decidedly poor science, and a setback for bread baking!  

Our generic lineup thus becomes (new named genera are bold italics):
Amelichloa
Ampelodesmos
Barkworthia
Eriocoma
Hesperostipa
Jarava
Nassella
Oloptum
Oryzopsis
Pappostipa
Piptatherum
Piptochaetium
Ptilagrostiella
Stipellula
Thorneochloa

The California roster: Format New Name             STATUS
Amelichloa brachychaeta (Godr.) Arriaga & Barkworth    Native
Amelichloa caudata (Trin.) Arriaga & Barkworth Waif
Amelichloa clandestina (Hack.) Arriaga & Barkworth         Waif
Ampelodesmos mauritanica (Poir.) T. Durand & Schinz,   Naturalized
Barkworthia stillmanii (Bol.) Romasch., P.M. Peterson & Soreng  Native
Eriocoma arida (M.E. Jones) Romasch.    Native
Eriocoma bloomeri (Bol.) Romasch.          Native
Eriocoma coronata (Thurb.) Romasch.    Native
Eriocoma hymenoides (Roem. & Schult.) Rydb.   Native
Eriocoma latiglumis (Swallen) Romasch. Native
Eriocoma lemmonii (Vasey) Romasch.    Native
Eriocoma lemmonii subsp. pubescens (Crampton) Romasch.        Native
Eriocoma lettermanii (Vasey) Romasch. Native
Eriocoma nelsonii (Scribn.) Romasch. subsp. dorei (Barkworth & J. Maze) Romasch.           Native
Eriocoma nevadensis (B.L. Johnson) Romasch.    Native
Eriocoma occidentalis  (Thurb. ex S. Watson) Romasch. subsp. californica (Merr. & Burtt Davy) Romasch. Native
Eriocoma occidentalis (Thurb. ex S. Watson) Romasch. ssp. occidentalis   Native
Eriocoma occidentalis (Thurb. ex S. Watson) Romasch. subsp. pubescens (Vasey) Romasch.,          Native
Eriocoma parishii (Vasey) Romasch.         Native
Eriocoma pinetorum (M.E. Jones) Romasch.         Native
Eriocoma thurberiana (Piper) Romasch. Native
Eriocoma webberi Thurb.             Native
Hesperostipa comata (Trin. & Rupr.) Barkworth subsp. comata    Native
Hesperostipa comata subsp. intermedia (Scribn. & Tweedy) Barkworth   Native
Jarava plumosa (Spreng.) S.W.L. Jacobs & J. Everett          Waif
Nassella cernua (Stebb. & Löve) Barkworth           Native
Nassella formicara (Del.) Barkworth         Native
Nassella lepida (A. Hitchc.) Barkworth     Native
Nassella manicata (E. Desv.) Barkworth  Naturalized
Nassella pulchra (A. Hitchc.) Barkworth  Native
Nassella tenuissima (Trin.) Barkworth     Waif
Nassella viridula (Trin.) Barkworth            Native
Oloptum miliaceum (L.) Röser & H.R. Hamasha   Naturalized
Oryzopsis exigua Thurb. Native
Oryzopsis micrantha (Trin. & Rupr.) Thurb.           Native
Pappostipa speciosa (Trin. & Rupr.) Romaschenko             Native
Piptatherum miliaceum (L.) Coss.              Naturalized
Piptochaetium setosum (Trin.) Arechav. Naturalized
Piptochaetium stipoides (Trin. & Rupr.) Hack. ex Arechav.              Naturalized
Ptilagrostiella kingii (Bol.) Romasch.         Native
Stipellula capensis (Thunb.) Röser & H.R. Hamasha            Naturalized
Thorneochloa diegoensis (Swallen) Romasch.      Native

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